It can be folded from either side. The upper left video folds from the left, the other from the right.
The results show that these characteristic asymmetrical patterns emerge as a result of simultaneous folding of overlapped wings. The revealed folding mechanisms can achieve not only highly compact wing storage but also immediate deployment. In addition, the right and left crease patterns are interchangeable, thus each wing internalizes two crease patterns and can be folded in two different ways. This two-way folding gives rove beetles freedom of choice for the folding direction. The use of asymmetric patterns and the capability of two-way folding are unique features not found in artificial structures. They have great potential to extend the design possibilities for all deployable structures, from space structures such as the solar panels on satellites to articles of daily use such as umbrellas and folding fans.